In the Southside Reef, a lowly bluestreak cleaner wrasse named Oscar fantasizes about being rich and famous. Soon after arriving for work at the Whale Wash, he is called to his boss's office, a pufferfish named Sykes, to discuss the fact that he owes a large sum of money and must pay it back by the next day. He remembers being humiliated as a child because his father was a tongue scrubber, so his angelfish best friend, Angie, offers him a shiny pink pearl that was a gift from her grandmother to pawn and pay his debt. Meanwhile, a gang of criminally-inclined sharks, with associates such as orcas, swordfish and octopuses, has a problem with Lenny, a vegetarian shark. His crime lord father, Don Edward Lino, orders his violent eldest son, Frankie, to mentor his brother in the family business.